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N65s were notorious for bad shutter release buttons. The metal is thinner than a strand of hair and they break. Search Ebay and see if the part is available to replace (note: Nikon stopped selling parts last year).
M90 is a battery independent setting - it requires no battery power to function. If it won't advance until put on M90 from other settings, then either you're not allowing for proper exposure, have the settings incorrectly set (film speed, etc), or you have shutter issues. The shutter is electronically controlled on all settings except M90. In the auto setting, your lense MUST be set to the smallest aperture (biggest number). On any other setting, you can use it however you'd like. I'd suggest unloading any film, setting the shutter speed dial to 1/125 or so, opening the back and looking thru and firing the shutter. If it's not snapping open and immediately closed, theres an issue and you need to consult a repair technician either locally, through Nikon, or through KEH Camera online.
Please follow these instructions:
1. Open the camera back
2. Pull the film leader across the red film index mark (red line)
3. Check to ensure film is properly positioned with no slack, then close camera back
4. Fully depress shutter release button to advance to frame 1. Check the rewind lever if it turns counter clockwise when you depress the shutter release button.
Keep on trying this procedure until you get it right. Most of the time, the error is in not pulling the film leader enough to reach the red film index mark.
If you're still having a problem with this, the next best option is to have the camera checked by a competent camera technician.
on page 101 (Troubleshooting) it mentions the Err message, and the shutter again to release it - According to the guide its a Refer Nikon Service Centre issue.
From how you describe it, it is consistent with there being a problem with lens compatibility. It would be worth checking compatibility (with the chart on pages 32 - 33) - Although if you inherited it "as a unit" I doubt thats the root cause..
Sounds like your batteries are near dead, enough juice to run the light and shutter control but not enough for the motor drive (which the control system knows it must use when film is loaded). The MB-21 lights not coming on is the battery test -- two lights means batteries OK, 1 light means low battery, no lights means exhausted.
Autofocus Lock ("AF-L"). Pressing the AF-L (Autofocus Lock) button enables the Nikon F4 to lock the focus. Simultaneous lock of AF-L and AE-L is possible. "'FREEZE FOCUS" - is another great function in the F4. It allows automatic shutter release when subject enters a pre focused distance - available with Nikon MF-23 and/or the 250 exp. MF-24 Multi-Control Back. i.e. with the shutter button fully depressed, the shutter is automatically fired when a subject comes into a preset manually focused position. This function is recommended for sports races because the racing course is usually known beforehand. It also works well in wildlife remote control photography, scientific, forensic and other technical photography.