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From the manual:
"If the error display is not canceled when the shutter release button is pressed again, insert the Dark slide, half-press the shutter release button while pressing the roll film holder's manual film advance button to wind up the film, then contact your nearest Mamiya agent or service center."
If you need a manual, you can download one from http://butkus.org/chinon/mamiya/mamiya_645af/mamiya_645af.htm
It sounds like the aux. curtain that covers the film when you're focusing either is not properly lowering, or is damaged somehow. You can remove the back and look through to see if you notice any light coming through. If you do, consult a tech. You'd only notice it on frames because mid-frames wouldn't remain in the light path long enough to be affected under most circumstances. It'd likely be a tear/nick in the curtain. Or, it could just not be completely be closing.
It could also be an issue with bad seals in the film back, or between the back and body, or even a crack somewhere in the back or the body itself. If you can't find a leak in the curtain, try keeping the dark slide in right up til you take a shot in, taking it out as soon as you're ready to shoot, replacing it before you wind, etc. to minimize the amount of time light could come from the anywhere but the back - to try and narrow down the location of the leak.
You might want to check and make sure the battery is still good. If you purchased the camera used and didn't change the battery then this might be the case.
But in the mean time this is what you need to do: 1st-make sure that the shutter speed dial is set to any shutter speed other than the red circle w/red ring. If your camera is set to that it will drain your battery when not in use since it is designed to be an automatic setting. I set mine to 1/125th to do this push in the circular dial on the side and at the same time turn the knob.
2nd- locate the 2 levers on the opposite side of the shutter speed dial. Once you found them then position the back lever all the way forward towards the setting for "MULTI" and position the lever towards the front straight up.
3rd- then wind the film crank till it won't wind anymore and depress the shutter, if it is wound and the shutter release is still stuck then skip to the fourth step, if something happened and you heard a click then still proceed to the next step because no the shutter release will be temporarily stuck.
4th-Once both of these levers are in this position there should be a red
button just behind the shutter release button on the top right side of
the camera.... drum roll please...."Push the red button" what you
should hear is a rather loud click, this click is the mirror releasing allowing you to: 1 wind the film crank and take another picture and 2 this also now allows you to see through the lens again (assuming that you have a lens on the camera).
In Conclusion: without a battery that is fully functional in the camera this is how the camera will operate once you get a battery for your camera then reposition the levers on the side of the camera opposite of what I suggested (this will be the normal operating mode for the camera with a battery) then when you wind the film crank and depress the shutter everything will function normally.
Easy to test with camera back open. Load a film and fire the shutter a few times each time with winding on, to see if film is winding correctly, at the same time you should be able notice if the shutters are firing, eight of them in succession makes a bit of noise.
This is not the best solution as it indicates a faulty film back. The film should advance without resorting to activating the film-wind release lever. That is only for intentionally advancing a partially exposed roll of film. Make sure the multi-exposure lever is not engaged and that the counter does advance and the red mark disappears as you advance to an unexposed frame.
You may not have loaded the film properly so that it wasn't wound forward after each exposure. Sometimes the film leader isn't picked up by the wind-on mechanism. Check the frame indicator moves as you wind the film on. Also operate the shutter without film and with the back open. You should see a flash of light through the lens if the shutter operates.
This is not a simple repair - if the wind lever is "ratcheting", parts will have to be replaced. If the wind lever is "free-wheeling", there might be no film in the camera, or the counter isn't working ( it has to be at #1 or above to cock the shutter ). It's a job for a repair shop.