When I am printing documents I often get black smudges on the right side of the paper. This happens often when I am printing an item with a lot of black ink (i.e. a clip art image that is mostly black). The smudges look like a duplication of items on the document, but placed lower on the document.
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It looks like you've managed to get some ink on the rollers. 3 Possible solutions:-
1) Check your printer maintenance menu, there may be a roller cleaning option.
2) Print a document which has printable data on almost all lines, but with empty cartridges. You'd be using a piece of paper more or less as a mop.
3) Final alternative, it getting right into your printer and cleaning it by hand. Printer ink is water soluble so damp cloths/cotton buds should be man enough for the job. The parts you need to concentrate on are the rollers and the small sponge wells underneath where the print cartridges rest.
The R2400 has an option to load paper on-top (Epson calls it the Automatic Document Feeder) and at the back. Based on the printer's User Guide the Velvet Fineart Paper should be loaded at the back. (Epson called is the Manual Single Sheet Feed).
Look at the User's Guide availabe on their website.
Very nice trouble shooting. The image defect you describe point to the OPC or drum unit this is essentially the toner cart. In your case black. Measureing the distance between repetitive defects is a sure way of diagnosing problems such as yours. Again nice work.
Google repetitive defect ruler for your model to help in the future.
Are you sure this is a Designjet 450 (large format printer) and not a Deskjet 450 (portable printer)? The following assumes you have wispy black streaking on the printed side of the paper. If the streaking is on the back side then post more details of the printer model and the issue.
In either case the most likely cause is a buildup of fibers on the underside of the printheads or carriage, causing wisps of black ink to be dragged on the paper when the carriage reverses direction. You can clean the printer as shown at the following (though the details will be somewhat different for the Designjet or Deskjet....).