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Both your air con and defogger (I am assuming you mean front windshield defroster setting, and not the back window) engage the air conditioner pump/compressor.
The screeching can be either a bad belt or a bad compressor.
How long has it been since the belt was changed? If it is old, and slipping, it will cause the screech at speed.
If the belt has recently been changed, and is adjusted properly, I would start looking that the air compressor pump.
on the left(driver's) side of the lower rad support. It's a small flat relay with a plug on the bottom and usually has a blue colored front. held on by two screws. Also, there may be more rad fan relays in the underhood fuse/relay box.